Gene T32N15.11
Putative Identification prephenate dehydratase
Position 87366 to 88640, from initial methionine to termination codon
Strand -
EST match T42435, N96431, H74930, N96696
Database match bacterial PheA proteins (prephenate dehydratase)

 

CDS:  The table below lists the coordinates of the single T32N15.11 exon and which exon prediction algorithms selected the termini (Gr = GRAIL)

Exon Range 3' 5'
1 87366 - 88640 Gr Gr

Complete CDS of T32N15.11

ATGCAAGCCGCAACGTCGTGTGATCTCAAGTTCCGATCAACAGATCCGACGTCTAGGAAC
AAATGTTTCTCCCACGCGATTCCAAAGCGCGTGGCTGTTACGTGCGGTTATAGGTCGGAG
TCGTTTAGCTTCCCTAACGGTGTCTCCGTGAGTCGATCTGATTGGCAAAGCTCATGTGCC
ATTTTATCTAGTAAAGTTGCTTCTGTTGAAAATACCGGTGGTTTAGCGGACAAAATCGCC
GCCGTTAATGGTCACACGAACGGCTCTGTGAATCTTGGTCTCGTCGCCGTTGAGTCAACT
AACGGAAAGTTAGCTCCTGCTCAGCCGTTGACTATTACTGATCTATCTCCGGCACCGTTG
CATGGTTCTAGTCTACGTGTAGCTTACCAAGGCGTTCCCGGAGCTTACTCGGAAGCAGCT
GCCGGAAAAGCTTATCCCAATTGCGACGCCATTCCTTGTGACCAGTTTGACGTTGCTTTT
CAGGCGGTGGAGCTTTGGATCGCTGATCGAGCTGTGCTTCCGGTGGAGAACTCACTCGGT
GGTTCGATCCATCGAAACTATGATCTTCTTCTCCGTCACCGTCTTCACATCGTTGGAGAA
GTTCAGATTCCGGTCCACCATTGCCTCCTCGCACTCCCGGGAGTCCGAACTGACTGTGTT
TCGCGGGTGATCTCTCACCCGCAAGCCCTAGCGCAGACGGAACACTCCCTCGACGTCCTT
ACACCACATGCAGCGCGTGAGGCTTTCCATGACACAGCGGCTGCTGCGGAGTATATCTCA
GCTAACGACCTACATGACACGGCGGCTGTGGCGAGCGCACGCGCCGCTGAGCTTTACAAC
CTCCAGATATTAGCGGATGGGATTCAAGACGACCCGGGGAACGTCACTCGTTTCCTAATG
TTGGCGCGTGAGCCTATAATCCCACGCACGGACCGGCCATTCAAGACTAGCATCGTCTTT
GCTGCCCAAGAGCACAAAGGAACTAGCGTCCTCTTTAAAGTCCTCTCGGCTTTCGCTTTC
AGAGACATTAGCCTGACCAAGATCGAGTCCCGGCCTCACCATAACCGCCCGCTTAGGGTT
GTTGGTGACGGGAGCTTTGGGACGTCGAAGAACTTCGAGTACATGTTTTACGTGGATTTT
GAGGCATCGATGGCTGAGCCGCGTGCACAGAACGCGCTTGCGGAGGTTCAAGAGTACACG
TCGTTCCTAAGGGTGCTGGGAAGTTACCCCATGGATATGACACCATGGTCCATGACATCC
ACAGAAGAAGCATGA

 

Protein translation:

MQAATSCDLKFRSTDPTSRNKCFSHAIPKRVAVTCGYRSESFSFPNGVSVSRSDWQSSCA
ILSSKVASVENTGGLADKIAAVNGHTNGSVNLGLVAVESTNGKLAPAQPLTITDLSPAPL
HGSSLRVAYQGVPGAYSEAAAGKAYPNCDAIPCDQFDVAFQAVELWIADRAVLPVENSLG
GSIHRNYDLLLRHRLHIVGEVQIPVHHCLLALPGVRTDCVSRVISHPQALAQTEHSLDVL
TPHAAREAFHDTAAAAEYISANDLHDTAAVASARAAELYNLQILADGIQDDPGNVTRFLM
LAREPIIPRTDRPFKTSIVFAAQEHKGTSVLFKVLSAFAFRDISLTKIESRPHHNRPLRV
VGDGSFGTSKNFEYMFYVDFEASMAEPRAQNALAEVQEYTSFLRVLGSYPMDMTPWSMTS
TEEA*

 

Biochemical reaction catalyzed prephanate dehydratase:

 COO-                          COO-                      COO-
 |                             |                         |
 C=O                           C=O                     H-C-NH3+
 |                             |                         |
 CH2                           CH2                       CH2
 |                             |                         |
(C6H5OH)-COO-                 (C6H5)                    (C6H5)
                PheA
Prephenate     ------->   Phenylpyruvate   ------->  Phenylalanine

In E. coli, the decarboxylation of prephanate to phenylpyruvate is catalyzed by prephenate dehydratase. Transamination of phenylpyruvate yields phenylalanine.

Alignment of bacterial PheA proteins and T32N15.11. The alignment contains the proteins encoded by the PheA genes of E. coli (M10431), E. herbicola (X60420), and P. stutzeri (M73971) with T32N15.11. The PROSITE Dictionary states that prephenate dehydratase (PDT, EC 4.2.1.51) catalyzes the decarboxylation of prephenate into phenylpyruvate. In microorganisms PDT is involved in the terminal pathway of the biosynthesis of phenylalanine. Signature sequences for prephenate dehydratase are centered around a conserved threonine which has been said to be essential for the activity of the enzyme in E. coli; the second region includes a conserved glutamate. These regions are highlighted in red.

            1                                                         60
 EcoliPheA  ~~~~~~~~~~MTSENPLLALREKISALDEKLLALLAERREL.....AVEVGKAKLLSHRP 
 EherbPheA  ~~~~~~~~~~MNPDNPLLALRDKISAVDKKLLTLLAERRLL.....AVEVAQAKLATHRP 
PstutzPheA  ~~~~~~~~~~MSEADQLKALRVRIDSLDERILDLISERARC.....AQEVARVKTASWPK 
 T32N15.11  MQAATSCDLKFRSTDPTSRNKCFSHAIPKRVAVTCGYRSESFSFPNGVSVSRSDWQSSCA 

            61                                                       120
 EcoliPheA  VRDI.....DRERDLLERLITLGKAHHLDAHYITRLFQLIIEDSVLTQQALLQQHLNKIN 
 EherbPheA  IRDV.....ERERALLENLIVLGKAHNLDAHYITRLFQLVIEDSVLTQQALLQKNLNHPH 
PstutzPheA  AEEAVFYRPEREAWVLKHIMELNKG.PLDNEEMARLFREIMSSCLALEQPL......... 
 T32N15.11  ILSSKVASVENTGGLADKIAAVN.GHTNGSVNLGLVAVESTNGKLAPAQPLTITDLSPAP 

            121                                                      180
 EcoliPheA  PH..SARIAFLGPKGSYSHLAARQYAARHFEQFIESGCAKFADIFNQVETGQADYAVVPI 
 EherbPheA  AH..AARIAFLGPKGSYSHLAARNYASRHFDSMVECGCLKFHDIIKQVENGVADYAVMPI 
PstutzPheA  ......RVAYLGPEGTFSQAAALKHFGH...SVISKPMAAIDEVFREVVAGAVNFGVVPV 
 T32N15.11  LHGSSLRVAYQGVPGAYSEAAA....GKAYPNCDAIPCDQFDVAFQAVELWIADRAVLPV 

            181                                                      240
 EcoliPheA  ENTSSGAINDVYDLLQHTSLSIVGEMTLTIDHCLLVSGTTDLSTINTVYSHPQPFQQCSK 
 EherbPheA  ENTSSGSINDVYDLLQQTSLSIVGELTLPIDHCVLVNGPTDLQQIETVYSHPQPFQQCSQ 
PstutzPheA  ENSTEGAVNHTLDSFLEHDIVICGEVELRIHHHLLVGETTKTDRITRIYSHAQSLAQCRK 
 T32N15.11  ENSLGGSIHRNYDLLLRHRLHIVGEVQIPVHHCLLALPGVRTDCVSRVISHPQALAQTEH 

            241                                                      300
 EcoliPheA  FLN.RYPHWKIEYTESTSAAMEKVAQAKSPHVAALGSEAGGTLYGLQVLERIEANQRQNF 
 EherbPheA  FIN.RFPHWKIEYTESTAAAMEKVAALNSPKVAALGSEAGGELYQLQVLERNLANQQQNH 
PstutzPheA  WLDAHYPNVERVAVSSNADAAKRVK..SEWNSAAIAGDMAAQLYGLSKLAEKIEDRPDNS 
 T32N15.11  SLDVLTPHAAREAFHDTAAAAEYISANDLHDTAAVASARAAELYNLQILADGIQDDPGNV 

            301                                                      360
 EcoliPheA  TRFVVLARKAINVSDQVPAKTTLLMATGQQAG..ALVEALLVLRNHNLIMTRLESRPIHG 
 EherbPheA  TRFIVLARKAIEVSDQVPAKTTLIMATGQQAG..ALVDALLVLRQHNLIMSKLESRPING 
PstutzPheA  TRFLIIGSQ..EVPPTGDDKTSIIVSMRNKPG..ALHELLMPFHSNGIDLTRIETRPSRS 
 T32N15.11  TRFLMLAREPIIPRTDRPFKTSIVFAAQEHKGTSVLFKVLSAFAFRDISLTKIESRPHHN 

            361                                                      420
 EcoliPheA  NP..............WEEMFYLDIQANLESAEMQKALKELGEITRSMKVLGCYPSENVV 
 EherbPheA  NP..............WEEMFYIDVQGNLQSERMQQALQELQTMTRSLKVLGCYPSENVV 
PstutzPheA  GK..............WTYVFFIDCMGHYQDPLIKDVLEKIDHEAVALKVLGSYPKAVL* 
 T32N15.11  RPLRVVGDGSFGTSKNFEYMFYVDFEASMAEPRAQNALAEVQEYTSFLRVLGSYPMDMTP 

            421  
 EcoliPheA  PVDPT*~~~~
 EherbPheA  PAEPGR*~~~
PstutzPheA  ~~~~~~~~~~
 T32N15.11  WSMTSTEEA*

 


written 28 Aug 97
Larry Parnell